The slant digging and down-raking action of hoofs on the steeper slopes of moraines has uprooted and buried many of the tender plants from year to year, without allowing them time to mature their seeds.

马蹄在冰碛的陡峭斜坡上的倾斜挖掘和向下耙动， 年复一年地连根拔起并掩埋了许多柔嫩的植物， 使它们的种子没有时间成熟。